While evolution is how life on earth came to be, where does the evolution occur? At what level of the hierarchy of life—from genes to individuals to groups to species—does evolution work its selection magic? Is it multilevel? It’s contentious.

What is the extended evolutionary synthesis (EES) and why better? Includes multilevel selection, transgenerational epigenetic inheritance, niche construction, evolvability, and ideas from evolutionary developmental biology. Why is EES controversial?